diff --git a/projects/pathway-browser/src/app/render/render-shapes.model.ts b/projects/pathway-browser/src/app/render/render-shapes.model.ts
index 208c38d..c4d87d2 100644
--- a/projects/pathway-browser/src/app/render/render-shapes.model.ts
+++ b/projects/pathway-browser/src/app/render/render-shapes.model.ts
@@ -51,6 +51,18 @@ export interface RenderNodeShape {
   dashed: boolean;
 }
 
+/**
+ * A polyline, and also the shape every arrowhead is drawn as.
+ *
+ * An arrowhead is geometry, not a line decoration. The diagram draws four of
+ * them and three carry meaning a triangle does not: catalysis is a hollow
+ * circle, positive regulation a hollow triangle, and negative regulation a bar
+ * across the line. OOXML's line ends offer `triangle`, `oval`, `stealth`,
+ * `diamond` and `arrow`, always filled and always the line's own colour -- so a
+ * tee has no spelling there at all, and an export that reached for the nearest
+ * line end drew inhibition as activation. Handing over the points instead means
+ * the exporter needs to know none of that.
+ */
 export interface RenderEdgeShape {
   kind: 'edge';
   id: string;
@@ -59,8 +71,9 @@ export interface RenderEdgeShape {
   points: { x: number; y: number }[];
   stroke: ShapeColour;
   strokeWidth: number;
-  /** Reactions draw arrowheads; a plain link does not. */
-  arrow: boolean;
+  /** Closed and filled, for an arrowhead; open and stroked, for a connector. */
+  closed: boolean;
+  fill: ShapeColour;
   /** A broken line, which the diagram uses for the links between glyphs. */
   dashed: boolean;
 }
diff --git a/projects/pathway-browser/src/app/render/render.component.ts b/projects/pathway-browser/src/app/render/render.component.ts
index 4dec3e6..aa7bab9 100644
--- a/projects/pathway-browser/src/app/render/render.component.ts
+++ b/projects/pathway-browser/src/app/render/render.component.ts
@@ -434,6 +434,21 @@ export class RenderComponent {
       !framed ||
       (box.x2 >= extent.x1 && box.x1 <= extent.x2 && box.y2 >= extent.y1 && box.y1 <= extent.y2);
 
+    // Keeping a glyph that merely crosses the frame is right; keeping the parts
+    // of it that fall outside is not. An SVG has a viewBox and simply does not
+    // draw them, and a slide has nothing of the sort -- a connector that ran the
+    // width of the diagram arrived as an object sitting 31 inches off the side
+    // of an 11 inch slide, which is what a reader would find when they opened it.
+    const clipBox = (box: { x1: number; y1: number; x2: number; y2: number }) =>
+      framed
+        ? {
+            x1: Math.max(box.x1, extent.x1),
+            y1: Math.max(box.y1, extent.y1),
+            x2: Math.min(box.x2, extent.x2),
+            y2: Math.min(box.y2, extent.y2),
+          }
+        : box;
+
     // toArray() rather than iterating the collection: a cytoscape collection is
     // array-like without being an array, and its filter hands back the loose
     // element type rather than a node.
@@ -448,9 +463,11 @@ export class RenderComponent {
     const nodeUnderlays: RenderNodeShape[] = [];
     const edges: RenderEdgeShape[] = [];
     const edgeUnderlays: RenderEdgeShape[] = [];
+    const arrowheads: (RenderNodeShape | RenderEdgeShape)[] = [];
 
     for (const node of visible) {
-      const box = node.boundingBox({ includeLabels: false, includeOverlays: false });
+      const box = clipBox(node.boundingBox({ includeLabels: false, includeOverlays: false }));
+      if (box.x2 <= box.x1 || box.y2 <= box.y1) continue;
       const label = String(node.data('displayName') ?? '').trim();
       const under = this.underlay(node);
       if (under) {
@@ -458,10 +475,14 @@ export class RenderComponent {
           kind: 'node',
           id: `${node.id()}-underlay`,
           name: `${label || node.id()} highlight`,
-          x: box.x1 - under.padding,
-          y: box.y1 - under.padding,
-          w: box.w + 2 * under.padding,
-          h: box.h + 2 * under.padding,
+          x: Math.max(box.x1 - under.padding, framed ? extent.x1 : -Infinity),
+          y: Math.max(box.y1 - under.padding, framed ? extent.y1 : -Infinity),
+          w:
+            Math.min(box.x2 + under.padding, framed ? extent.x2 : Infinity) -
+            Math.max(box.x1 - under.padding, framed ? extent.x1 : -Infinity),
+          h:
+            Math.min(box.y2 + under.padding, framed ? extent.y2 : Infinity) -
+            Math.max(box.y1 - under.padding, framed ? extent.y1 : -Infinity),
           geom: this.geometryOf(String(node.style('shape') ?? '')),
           fill: under.colour,
           stroke: null,
@@ -480,8 +501,8 @@ export class RenderComponent {
         name: label || String(node.data('schemaClass') ?? node.id()),
         x: box.x1,
         y: box.y1,
-        w: box.w,
-        h: box.h,
+        w: box.x2 - box.x1,
+        h: box.y2 - box.y1,
         geom: this.geometryOf(String(node.style('shape') ?? '')),
         fill: this.colour(node.style('background-color'), node.style('background-opacity')),
         stroke: this.colour(node.style('border-color'), node.style('border-opacity')),
@@ -501,34 +522,58 @@ export class RenderComponent {
       .filter((edge) => within(edge.boundingBox()))) {
       // Bend points are how the diagram's orthogonal routing is stored; without
       // them every connector exports as a straight diagonal.
-      const points = [edge.sourceEndpoint(), ...this.bendPoints(edge), edge.targetEndpoint()];
+      const whole = [edge.sourceEndpoint(), ...this.bendPoints(edge), edge.targetEndpoint()].map(
+        (point) => ({ x: point.x, y: point.y })
+      );
+      // Unframed, this is the polyline itself; framed, only the parts in view.
+      const runs = framed ? this.clipRuns(whole, extent) : [whole];
+      if (!runs.length) continue;
+      const points = whole;
       const width = this.pixels(edge.style('width')) || 1;
       const under = this.underlay(edge);
       if (under) {
-        edgeUnderlays.push({
-          kind: 'edge',
-          id: `${edge.id()}-underlay`,
-          name: `${String(edge.data('schemaClass') ?? 'connector')} highlight`,
-          points: points.map((point) => ({ x: point.x, y: point.y })),
-          stroke: under.colour,
-          // Cytoscape pads an underlay outwards from the line, so the band is
-          // the line plus that padding on each side.
-          strokeWidth: width + 2 * under.padding,
-          arrow: false,
-          dashed: false,
+        // One band per run, or a connector clipped into two pieces keeps its
+        // tint on the first and loses it on the rest.
+        runs.forEach((run, at) => {
+          edgeUnderlays.push({
+            kind: 'edge',
+            id: at === 0 ? `${edge.id()}-underlay` : `${edge.id()}-underlay-${at}`,
+            name: `${String(edge.data('schemaClass') ?? 'connector')} highlight`,
+            points: run,
+            stroke: under.colour,
+            // Cytoscape pads an underlay outwards from the line, so the band is
+            // the line plus that padding on each side.
+            strokeWidth: width + 2 * under.padding,
+            closed: false,
+            fill: null,
+            dashed: false,
+          });
         });
       }
 
-      edges.push({
-        kind: 'edge',
-        id: String(edge.id()),
-        name: String(edge.data('schemaClass') ?? 'connector'),
-        points: points.map((point) => ({ x: point.x, y: point.y })),
-        stroke: this.colour(edge.style('line-color'), edge.style('opacity')),
-        strokeWidth: width,
-        arrow: String(edge.style('target-arrow-shape') ?? 'none') !== 'none',
-        dashed: this.dashed(edge.style('line-style')),
+      const stroke = this.colour(edge.style('line-color'), edge.style('opacity'));
+      const dashed = this.dashed(edge.style('line-style'));
+      runs.forEach((run, at) => {
+        edges.push({
+          kind: 'edge',
+          id: at === 0 ? String(edge.id()) : `${edge.id()}-${at}`,
+          name: String(edge.data('schemaClass') ?? 'connector'),
+          points: run,
+          stroke,
+          strokeWidth: width,
+          closed: false,
+          fill: null,
+          dashed,
+        });
       });
+
+      // The arrowhead is drawn after its line, so it sits on top of it -- and
+      // only when the end it marks is in view.
+      const tip = points[points.length - 1];
+      const showsTip =
+        !framed ||
+        (tip.x >= extent.x1 && tip.x <= extent.x2 && tip.y >= extent.y1 && tip.y <= extent.y2);
+      if (showsTip) arrowheads.push(...this.arrowhead(edge, points, width));
     }
 
     return {
@@ -544,7 +589,14 @@ export class RenderComponent {
       // The underlays are how the diagram tints an event by the sub-pathway it
       // belongs to, and each sits immediately behind the glyph it marks -- so
       // they go in front of the compartments and behind everything else.
-      shapes: [...compartments, ...edgeUnderlays, ...edges, ...nodeUnderlays, ...nodes],
+      shapes: [
+        ...compartments,
+        ...edgeUnderlays,
+        ...edges,
+        ...arrowheads,
+        ...nodeUnderlays,
+        ...nodes,
+      ],
     };
   }
 
@@ -625,6 +677,185 @@ export class RenderComponent {
     return null;
   }
 
+  /**
+   * A polyline cut down to a rectangle, as the runs that survive.
+   *
+   * Liang-Barsky per segment. A connector crossing the frame twice comes back
+   * as two runs rather than one line joining them through the middle, which is
+   * what naive clipping to the endpoints would draw.
+   */
+  private clipRuns(
+    points: { x: number; y: number }[],
+    box: { x1: number; y1: number; x2: number; y2: number }
+  ): { x: number; y: number }[][] {
+    const runs: { x: number; y: number }[][] = [];
+    let run: { x: number; y: number }[] = [];
+
+    const near = (a: { x: number; y: number }, b: { x: number; y: number }) =>
+      Math.abs(a.x - b.x) < 0.01 && Math.abs(a.y - b.y) < 0.01;
+
+    for (let at = 0; at < points.length - 1; at++) {
+      const from = points[at];
+      const to = points[at + 1];
+      const dx = to.x - from.x;
+      const dy = to.y - from.y;
+      let enter = 0;
+      let leave = 1;
+      let inside = true;
+
+      for (const [p, q] of [
+        [-dx, from.x - box.x1],
+        [dx, box.x2 - from.x],
+        [-dy, from.y - box.y1],
+        [dy, box.y2 - from.y],
+      ]) {
+        if (p === 0) {
+          // Parallel to this edge: outside it means the whole segment is out.
+          if (q < 0) {
+            inside = false;
+            break;
+          }
+          continue;
+        }
+        const at2 = q / p;
+        if (p < 0) enter = Math.max(enter, at2);
+        else leave = Math.min(leave, at2);
+      }
+      if (!inside || enter > leave) {
+        if (run.length > 1) runs.push(run);
+        run = [];
+        continue;
+      }
+
+      const start = { x: from.x + dx * enter, y: from.y + dy * enter };
+      const end = { x: from.x + dx * leave, y: from.y + dy * leave };
+      if (!run.length) run.push(start);
+      else if (!near(run[run.length - 1], start)) {
+        if (run.length > 1) runs.push(run);
+        run = [start];
+      }
+      run.push(end);
+      // A segment that left the box ends the run.
+      if (leave < 1) {
+        runs.push(run);
+        run = [];
+      }
+    }
+
+    if (run.length > 1) runs.push(run);
+    return runs;
+  }
+
+  /**
+   * The mark at the target end of a connector, as geometry.
+   *
+   * The diagram uses four, and three of them mean something a triangle does
+   * not: catalysis is a hollow circle, positive regulation a hollow triangle,
+   * and negative regulation a bar across the line. Exporting them all as a
+   * filled triangle drew inhibition as activation, which is a figure that says
+   * the opposite of what the pathway does.
+   *
+   * Built here rather than in the exporter because it is diagram geometry --
+   * where the line ends, which way it points -- and because OOXML's own line
+   * ends cannot express a bar, a hollow head, or a colour of their own.
+   */
+  private arrowhead(
+    edge: cytoscape.EdgeSingular,
+    points: { x: number; y: number }[],
+    width: number
+  ): (RenderNodeShape | RenderEdgeShape)[] {
+    const shape = String(edge.style('target-arrow-shape') ?? 'none');
+    if (shape === 'none' || points.length < 2) return [];
+
+    const colour = this.colour(edge.style('target-arrow-color'), edge.style('opacity'));
+    if (!colour) return [];
+
+    const hollow = String(edge.style('target-arrow-fill') ?? 'filled') === 'hollow';
+    // Cytoscape scales an arrow by the line's width; measured against the
+    // exported SVG, a 4px connector carries a 16px head.
+    const scale = this.pixels(edge.style('arrow-scale')) || 1;
+    const size = Math.max(6, width * 4 * scale);
+
+    const tip = points[points.length - 1];
+    const from = points[points.length - 2];
+    const run = Math.hypot(tip.x - from.x, tip.y - from.y) || 1;
+    // Along the line, and across it.
+    const ax = (tip.x - from.x) / run;
+    const ay = (tip.y - from.y) / run;
+    const cx = -ay;
+    const cy = ax;
+
+    const name = `${String(edge.data('schemaClass') ?? 'connector')} ${shape}`;
+    const id = `${edge.id()}-arrow`;
+
+    if (shape === 'tee') {
+      // A bar across the line's end. Its thickness is the line's, not the
+      // head's, or inhibition on a hairline connector becomes a black square.
+      return [
+        {
+          kind: 'edge',
+          id,
+          name,
+          points: [
+            { x: tip.x + cx * (size / 2), y: tip.y + cy * (size / 2) },
+            { x: tip.x - cx * (size / 2), y: tip.y - cy * (size / 2) },
+          ],
+          stroke: colour,
+          strokeWidth: Math.max(width, size / 4),
+          closed: false,
+          fill: null,
+          dashed: false,
+        },
+      ];
+    }
+
+    if (shape === 'circle') {
+      // Centred where the line stops, which is how cytoscape seats it.
+      return [
+        {
+          kind: 'node',
+          id,
+          name,
+          x: tip.x - ax * (size / 2) - size / 2,
+          y: tip.y - ay * (size / 2) - size / 2,
+          w: size,
+          h: size,
+          geom: 'ellipse',
+          fill: hollow ? null : colour,
+          stroke: colour,
+          strokeWidth: hollow ? Math.max(1, width) : 0,
+          label: '',
+          fontSize: 0,
+          fontColor: null,
+          bold: false,
+          dashed: false,
+        },
+      ];
+    }
+
+    // Everything else is a triangle: the tip on the line's end, the base back
+    // along it. `stealth`, `diamond` and the rest do not occur in this diagram,
+    // and a triangle is the honest approximation if one ever does.
+    const base = { x: tip.x - ax * size, y: tip.y - ay * size };
+    return [
+      {
+        kind: 'edge',
+        id,
+        name,
+        points: [
+          { x: tip.x, y: tip.y },
+          { x: base.x + cx * (size / 2), y: base.y + cy * (size / 2) },
+          { x: base.x - cx * (size / 2), y: base.y - cy * (size / 2) },
+        ],
+        stroke: colour,
+        strokeWidth: hollow ? Math.max(1, width) : 0,
+        closed: true,
+        fill: hollow ? null : colour,
+        dashed: false,
+      },
+    ];
+  }
+
   /**
    * The band the diagram draws behind a glyph, if it draws one.
    *
diff --git a/tools/render/README.md b/tools/render/README.md
index a771927..452b591 100644
--- a/tools/render/README.md
+++ b/tools/render/README.md
@@ -82,6 +82,12 @@ diagram looks like. R-HSA-109606 comes out as **635 shapes in 32 KB**, against
 The slide is the size of the diagram, as production's is, so labels land at 5–53pt
 rather than the 1.65pt that fitting a 5976px diagram onto a 13.3in slide gives.
 
+Arrowheads are geometry rather than OOXML line ends. The diagram draws four and
+three of them mean something a triangle does not — catalysis is a hollow circle,
+positive regulation a hollow triangle, negative regulation a bar across the line
+— while a line end is always filled and always the line's own colour, and has no
+spelling for a bar at all. Drawn as line ends, inhibition came out as activation.
+
 Two things a slide of shapes does not carry yet:
 
 - **Node decorations.** The style draws them with `background-image` — 140 small
diff --git a/tools/render/fixtures/shapes-apoptosis.json b/tools/render/fixtures/shapes-apoptosis.json
index f080086..0655b49 100644
--- a/tools/render/fixtures/shapes-apoptosis.json
+++ b/tools/render/fixtures/shapes-apoptosis.json
@@ -132,7 +132,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -155,7 +156,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -178,7 +180,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -197,7 +200,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -216,7 +220,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -239,7 +244,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -262,7 +268,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -285,7 +292,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -308,7 +316,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -331,7 +340,8 @@
    ],
    "stroke": "0000cc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -350,7 +360,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -373,7 +384,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -396,7 +408,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -419,7 +432,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -438,7 +452,8 @@
    ],
    "stroke": "cccc0024",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -457,7 +472,8 @@
    ],
    "stroke": "cccc0024",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -476,7 +492,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -499,7 +516,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -518,7 +536,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -541,7 +560,8 @@
    ],
    "stroke": "cccc0024",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -560,7 +580,8 @@
    ],
    "stroke": "cccc0024",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -583,7 +604,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -602,7 +624,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -633,7 +656,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -656,7 +680,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -687,7 +712,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -710,7 +736,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -733,7 +760,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -760,7 +788,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -787,7 +816,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -810,7 +840,8 @@
    ],
    "stroke": "00cccc24",
    "strokeWidth": 44,
-   "arrow": false,
+   "closed": false,
+   "fill": null,
    "dashed": false
   },
   {
@@ -833,7 +864,8 @@
    ],
    "stroke": "00cccc24",
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diff --git a/tools/render/fixtures/shapes-framed-reaction.json b/tools/render/fixtures/shapes-framed-reaction.json
new file mode 100644
index 0000000..a3f3e67
--- /dev/null
+++ b/tools/render/fixtures/shapes-framed-reaction.json
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+    {
+     "x": 3932,
+     "y": 1430
+    },
+    {
+     "x": 3956.199914079815,
+     "y": 1418.4084445163912
+    },
+    {
+     "x": 3955.7931928347757,
+     "y": 1442.4049979736865
+    }
+   ],
+   "stroke": "001f25",
+   "strokeWidth": 0,
+   "closed": true,
+   "fill": "001f25",
+   "dashed": false
+  },
+  {
+   "kind": "edge",
+   "id": "3678-arrow",
+   "name": "connector triangle",
+   "points": [
+    {
+     "x": 3902,
+     "y": 1360
+    },
+    {
+     "x": 3927.800394907026,
+     "y": 1352.6284585979924
+    },
+    {
+     "x": 3923.3774700658214,
+     "y": 1376.2173910844162
+    }
+   ],
+   "stroke": "001f25",
+   "strokeWidth": 0,
+   "closed": true,
+   "fill": "001f25",
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2851",
+   "name": "2851",
+   "x": 3971,
+   "y": 1241,
+   "w": 30,
+   "h": 30,
+   "geom": "ellipse",
+   "fill": "001f25",
+   "stroke": "001f25",
+   "strokeWidth": 4,
+   "label": "",
+   "fontSize": 16,
+   "fontColor": "001f25",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2981",
+   "name": "2981",
+   "x": 4175,
+   "y": 1305,
+   "w": 30,
+   "h": 30,
+   "geom": "ellipse",
+   "fill": "001f25",
+   "stroke": "001f25",
+   "strokeWidth": 4,
+   "label": "",
+   "fontSize": 16,
+   "fontColor": "001f25",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2144",
+   "name": "2144",
+   "x": 3748,
+   "y": 1560,
+   "w": 32,
+   "h": 32,
+   "geom": "roundRect",
+   "fill": "effbff",
+   "stroke": "0561a6",
+   "strokeWidth": 6,
+   "label": "",
+   "fontSize": 16,
+   "fontColor": "001f25",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2823",
+   "name": "XIAP",
+   "x": 4151,
+   "y": 1131,
+   "w": 74,
+   "h": 46,
+   "geom": "roundRect",
+   "fill": "001f29",
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "XIAP",
+   "fontSize": 6,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "3374",
+   "name": "GSDMD(1-​275):​cardiolipin",
+   "x": 3722,
+   "y": 1393,
+   "w": 204,
+   "h": 74,
+   "geom": "rect",
+   "fill": null,
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "GSDMD(1-​275):​cardiolipin",
+   "fontSize": 12,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "3375",
+   "name": "GSDME:​cardiolipin",
+   "x": 3660,
+   "y": 1319,
+   "w": 236,
+   "h": 46,
+   "geom": "rect",
+   "fill": null,
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "GSDME:​cardiolipin",
+   "fontSize": 12,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "3674",
+   "name": "Pyroptosis",
+   "x": 4054,
+   "y": 1394,
+   "w": 176,
+   "h": 84,
+   "geom": "rect",
+   "fill": "006782",
+   "stroke": "ffffff",
+   "strokeWidth": 12,
+   "label": "Pyroptosis",
+   "fontSize": 12,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2145",
+   "name": "CYCS",
+   "x": 3599,
+   "y": 1555,
+   "w": 82,
+   "h": 46,
+   "geom": "roundRect",
+   "fill": "001f29",
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "CYCS",
+   "fontSize": 6,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2146",
+   "name": "CYCS",
+   "x": 3877,
+   "y": 1555,
+   "w": 82,
+   "h": 46,
+   "geom": "roundRect",
+   "fill": "001f29",
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "CYCS",
+   "fontSize": 6,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2982",
+   "name": "SEPT4 dimer",
+   "x": 3914,
+   "y": 1301,
+   "w": 176,
+   "h": 46,
+   "geom": "rect",
+   "fill": null,
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "SEPT4 dimer",
+   "fontSize": 12,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2496",
+   "name": "CASP3(1-​277) dimer",
+   "x": 4214,
+   "y": 992.1363636363637,
+   "w": 20.181818181818016,
+   "h": 60.86363636363626,
+   "geom": "rect",
+   "fill": null,
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "CASP3(1-​277) dimer",
+   "fontSize": 12,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  },
+  {
+   "kind": "node",
+   "id": "2531",
+   "name": "Activated BAX,​BAK oligomers",
+   "x": 3473,
+   "y": 1031,
+   "w": 270,
+   "h": 90,
+   "geom": "roundRect",
+   "fill": null,
+   "stroke": "000000",
+   "strokeWidth": 0,
+   "label": "Activated BAX,​BAK oligomers",
+   "fontSize": 12,
+   "fontColor": "ffffff",
+   "bold": false,
+   "dashed": false
+  }
+ ]
+}
diff --git a/tools/render/pptx.mjs b/tools/render/pptx.mjs
index beb3821..3a5ac87 100644
--- a/tools/render/pptx.mjs
+++ b/tools/render/pptx.mjs
@@ -258,7 +258,11 @@ function textBody({ label, fontSize, fontColor, bold, emuPerPx }) {
   return (
     `<p:txBody>${body}<a:p><a:pPr algn="ctr"/><a:r>` +
     `<a:rPr lang="en-US" sz="${points}"${bold ? ' b="1"' : ''} dirty="0">` +
-    (colour ? `<a:solidFill><a:srgbClr val="${colour.hex}"/></a:solidFill>` : '') +
+    (colour
+      ? `<a:solidFill><a:srgbClr val="${colour.hex}">` +
+        (colour.alpha < 1 ? `<a:alpha val="${Math.round(colour.alpha * 100000)}"/>` : '') +
+        `</a:srgbClr></a:solidFill>`
+      : '') +
     `</a:rPr><a:t>${escapeXml(label)}</a:t></a:r></a:p></p:txBody>`
   );
 }
@@ -266,13 +270,17 @@ function textBody({ label, fontSize, fontColor, bold, emuPerPx }) {
 /** A node: a preset rectangle, rounded rectangle or ellipse, with its label. */
 function nodeShape(shape, id, place, emuPerPx) {
   const at = place(shape.x, shape.y, shape.w, shape.h);
+  const stroke =
+    shape.strokeWidth > 0 || !shape.fill
+      ? outline(shape.stroke, shape.strokeWidth, emuPerPx, { dashed: Boolean(shape.dashed) })
+      : '<a:ln><a:noFill/></a:ln>';
   return (
     `<p:sp><p:nvSpPr><p:cNvPr id="${id}" name="${escapeXml(shape.name || shape.id)}"/>` +
     `<p:cNvSpPr/><p:nvPr/></p:nvSpPr><p:spPr>` +
     `<a:xfrm><a:off x="${at.x}" y="${at.y}"/><a:ext cx="${at.cx}" cy="${at.cy}"/></a:xfrm>` +
     `<a:prstGeom prst="${shape.geom}"><a:avLst/></a:prstGeom>` +
     solidFill(shape.fill) +
-    outline(shape.stroke, shape.strokeWidth, emuPerPx, { dashed: Boolean(shape.dashed) }) +
+    stroke +
     `</p:spPr>` +
     textBody({ ...shape, emuPerPx }) +
     `</p:sp>`
@@ -290,6 +298,8 @@ function nodeShape(shape, id, place, emuPerPx) {
  * is legal but which PowerPoint handles badly, so both sides have a floor.
  */
 function edgeShape(shape, id, place, emuPerPx) {
+  // Closed and filled means an arrowhead; the page hands those over as points
+  // for the same reason it hands over everything else.
   const points = (shape.points ?? []).filter(
     (point) => Number.isFinite(point?.x) && Number.isFinite(point?.y)
   );
@@ -302,14 +312,15 @@ function edgeShape(shape, id, place, emuPerPx) {
   const cx = Math.max(1, at.cx);
   const cy = Math.max(1, at.cy);
 
-  const path = points
-    .map((point, index) => {
-      const x = Math.round((point.x - box.x) * emuPerPx);
-      const y = Math.round((point.y - box.y) * emuPerPx);
-      const pt = `<a:pt x="${x}" y="${y}"/>`;
-      return index === 0 ? `<a:moveTo>${pt}</a:moveTo>` : `<a:lnTo>${pt}</a:lnTo>`;
-    })
-    .join('');
+  const path =
+    points
+      .map((point, index) => {
+        const x = Math.round((point.x - box.x) * emuPerPx);
+        const y = Math.round((point.y - box.y) * emuPerPx);
+        const pt = `<a:pt x="${x}" y="${y}"/>`;
+        return index === 0 ? `<a:moveTo>${pt}</a:moveTo>` : `<a:lnTo>${pt}</a:lnTo>`;
+      })
+      .join('') + (shape.closed ? '<a:close/>' : '');
 
   return (
     `<p:sp><p:nvSpPr><p:cNvPr id="${id}" name="${escapeXml(shape.name || shape.id)}"/>` +
@@ -318,11 +329,13 @@ function edgeShape(shape, id, place, emuPerPx) {
     `<a:custGeom><a:avLst/><a:gdLst/><a:ahLst/><a:cxnLst/>` +
     `<a:rect l="0" t="0" r="r" b="b"/><a:pathLst>` +
     `<a:path w="${cx}" h="${cy}">${path}</a:path>` +
-    `</a:pathLst></a:custGeom><a:noFill/>` +
-    outline(shape.stroke, shape.strokeWidth, emuPerPx, {
-      dashed: Boolean(shape.dashed),
-      extra: shape.arrow ? '<a:tailEnd type="triangle" w="med" len="med"/>' : '',
-    }) +
+    `</a:pathLst></a:custGeom>` +
+    solidFill(shape.fill) +
+    // A filled arrowhead has no outline to draw; giving it one at width 0 makes
+    // PowerPoint draw a hairline, which on a small head is most of the head.
+    (shape.strokeWidth > 0 || !shape.fill
+      ? outline(shape.stroke, shape.strokeWidth, emuPerPx, { dashed: Boolean(shape.dashed) })
+      : '<a:ln><a:noFill/></a:ln>') +
     `</p:spPr><p:txBody><a:bodyPr/><a:lstStyle/><a:p/></p:txBody></p:sp>`
   );
 }
diff --git a/tools/render/pptx.spec.mjs b/tools/render/pptx.spec.mjs
index fbe7ffe..acf39a4 100644
--- a/tools/render/pptx.spec.mjs
+++ b/tools/render/pptx.spec.mjs
@@ -96,7 +96,25 @@ const EDGE = {
   ],
   stroke: 'rgb(0,0,0)',
   strokeWidth: 1.2,
-  arrow: true,
+  closed: false,
+  fill: null,
+  dashed: false,
+};
+
+/** An arrowhead, which the page hands over as a closed, filled polygon. */
+const ARROW = {
+  kind: 'edge',
+  id: 'e1-arrow',
+  name: 'production triangle',
+  points: [
+    { x: 60, y: 20 },
+    { x: 48, y: 26 },
+    { x: 48, y: 14 },
+  ],
+  stroke: '001f25',
+  strokeWidth: 0,
+  closed: true,
+  fill: '001f25',
   dashed: false,
 };
 
@@ -207,13 +225,40 @@ describe('a slide built from shapes', () => {
     expect((slide.match(/<a:moveTo>/g) ?? []).length).toBe(1);
   });
 
-  it('gives a reaction an arrowhead and a plain link none', () => {
-    const arrow = open(pptx({ shapes: { ...SHAPES, shapes: [EDGE] } })).slide();
-    const plain = open(
-      pptx({ shapes: { ...SHAPES, shapes: [{ ...EDGE, arrow: false }] } })
+  it('closes and fills an arrowhead, and leaves a connector open', () => {
+    // OOXML line ends are always filled and always the line's own colour, so an
+    // arrowhead drawn as one cannot be hollow, cannot be a bar, and cannot be
+    // green. The page sends geometry instead; this is the shape of that.
+    const head = open(pptx({ shapes: { ...SHAPES, shapes: [ARROW] } })).slide();
+    expect(head).toContain('<a:close/>');
+    expect(head).toContain('<a:srgbClr val="001F25">');
+    expect(head, 'a filled head draws no outline of its own').toContain('<a:ln><a:noFill/></a:ln>');
+
+    const line = open(pptx({ shapes: { ...SHAPES, shapes: [EDGE] } })).slide();
+    expect(line).not.toContain('<a:close/>');
+    expect(line).toContain('<a:noFill/>');
+  });
+
+  it('leaves a hollow arrowhead unfilled and outlined', () => {
+    const slide = open(
+      pptx({
+        shapes: {
+          ...SHAPES,
+          shapes: [{ ...ARROW, fill: null, stroke: '0c9509', strokeWidth: 2 }],
+        },
+      })
     ).slide();
-    expect(arrow).toContain('tailEnd');
-    expect(plain).not.toContain('tailEnd');
+    expect(slide).toContain('<a:close/>');
+    expect(slide).toContain('<a:noFill/>');
+    expect(slide, 'the outline carries the arrowhead colour').toContain('<a:srgbClr val="0C9509">');
+  });
+
+  it('never spells an arrowhead as a line end', () => {
+    // A line end would be the obvious way to draw one and cannot say what the
+    // diagram says: a tee has no line end at all.
+    const slide = open(pptx({ shapes: REAL })).slide();
+    expect(slide).not.toContain('tailEnd');
+    expect(slide).not.toContain('headEnd');
   });
 
   it('keeps the diagram square, mapping both axes by one scale', () => {
@@ -376,10 +421,12 @@ describe('a slide built from a payload the page produced', () => {
         expect(slide, fill).toContain(`<a:alpha val="${permille}"/>`);
       }
     }
-    // And nothing the page coloured may come out unfilled: one noFill per
-    // uncoloured node, plus one per edge, and no more.
-    const uncoloured = REAL.shapes.filter((shape) => shape.kind === 'edge' || !shape.fill).length;
-    expect((slide.match(/<a:noFill\/>/g) ?? []).length).toBe(uncoloured);
+    // And nothing the page coloured may come out unfilled. An outline can be
+    // noFill too, so strip the outlines first: what is left is one body fill
+    // per shape, empty exactly where the page sent no colour.
+    const bodies = slide.replace(/<a:ln[ >][\s\S]*?<\/a:ln>/g, '');
+    const unfilled = REAL.shapes.filter((shape) => !shape.fill).length;
+    expect((bodies.match(/<a:noFill\/>/g) ?? []).length).toBe(unfilled);
   });
 
   it('keeps every label the page gave it, escaped', () => {
@@ -479,6 +526,68 @@ describe('the slide itself', () => {
   });
 });
 
+describe('a payload framed on one event', () => {
+  /**
+   * The same diagram asked for one reaction, which is what `?select=` does.
+   *
+   * Here because framing is where the contract "every shape lies inside the
+   * extent" is easy to break and impossible to notice: an SVG has a viewBox and
+   * simply does not draw what falls outside, so the same payload looks right in
+   * one format and puts objects 31 inches off the side of the slide in another.
+   */
+  const FRAMED = JSON.parse(
+    readFileSync(new URL('./fixtures/shapes-framed-reaction.json', import.meta.url), 'utf8')
+  );
+
+  it('is a frame, not the whole diagram', () => {
+    expect(FRAMED.shapes.length).toBeLessThan(REAL.shapes.length / 4);
+    expect(FRAMED.width).toBeLessThan(REAL.width / 2);
+  });
+
+  it.each([
+    ['framed', () => FRAMED],
+    ['whole', () => REAL],
+  ])('keeps every %s shape inside the extent it declares', (_name, payload) => {
+    const page = payload();
+    const right = page.x + page.width;
+    const bottom = page.y + page.height;
+    // A hair of tolerance: the extent and the geometry are both floats.
+    const slack = 0.5;
+    for (const shape of page.shapes) {
+      const points =
+        shape.kind === 'edge'
+          ? shape.points
+          : [
+              { x: shape.x, y: shape.y },
+              { x: shape.x + shape.w, y: shape.y + shape.h },
+            ];
+      for (const point of points) {
+        expect(point.x, `${shape.name} x`).toBeGreaterThanOrEqual(page.x - slack);
+        expect(point.x, `${shape.name} x`).toBeLessThanOrEqual(right + slack);
+        expect(point.y, `${shape.name} y`).toBeGreaterThanOrEqual(page.y - slack);
+        expect(point.y, `${shape.name} y`).toBeLessThanOrEqual(bottom + slack);
+      }
+    }
+  });
+
+  it('puts no shape past the edge of the slide it is drawn on', () => {
+    const { files, drawn } = open(pptx({ title: 'Framed', shapes: FRAMED }));
+    const [, cx, cy] = /<p:sldSz cx="(\d+)" cy="(\d+)"\/>/.exec(
+      strFromU8(files['ppt/presentation.xml'])
+    );
+    const placed = [
+      ...drawn().matchAll(/<a:off x="(-?\d+)" y="(-?\d+)"\/><a:ext cx="(\d+)" cy="(\d+)"\/>/g),
+    ];
+    expect(placed.length).toBeGreaterThan(10);
+    for (const [, x, y, w, h] of placed) {
+      expect(Number(x)).toBeGreaterThanOrEqual(0);
+      expect(Number(y)).toBeGreaterThanOrEqual(0);
+      expect(Number(x) + Number(w)).toBeLessThanOrEqual(Number(cx));
+      expect(Number(y) + Number(h)).toBeLessThanOrEqual(Number(cy));
+    }
+  });
+});
+
 describe('the tints the diagram paints behind its glyphs', () => {
   it('draws every underlay the page described', () => {
     const underlays = REAL.shapes.filter((shape) => shape.name.endsWith('highlight'));
diff --git a/tools/render/service.mjs b/tools/render/service.mjs
index 6f8e3bf..fec8739 100644
--- a/tools/render/service.mjs
+++ b/tools/render/service.mjs
@@ -63,8 +63,15 @@ const CACHE = process.env.RENDER_CACHE || path.resolve('.render-cache');
 // does: it keys the disk cache AND is the ETag, so it is the only thing that
 // tells a browser its copy is stale. v2 = full-size differenced GIFs; v3 =
 // illustrations export as standalone SVG, styles inlined and size written down;
-// v4 = PowerPoint is shapes rather than a picture of them.
-const CACHE_KEY = process.env.RENDER_CACHE_KEY || 'v4';
+// v4 = PowerPoint is shapes rather than a picture of them; v5 = arrowheads are
+// geometry, so a hollow circle and an inhibition bar are no longer both drawn
+// as a filled triangle.
+//
+// v5 exists because v4 was already published and the exporter changed again --
+// the second time in one day that a correct build served a stale file. If you
+// changed anything under tools/render that affects output, this line is part of
+// the change.
+const CACHE_KEY = process.env.RENDER_CACHE_KEY || 'v5';
 const CONCURRENCY = Number(process.env.RENDER_CONCURRENCY || 2);
 // Generous next to a real render, which is 3-8s, but far short of the two
 // minutes a page that never becomes ready would otherwise hold a browser for.